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Robert Thorn (Liev Schreiber), an American diplomat stationed in Italy, is told that his first-born son died soon after birth and to spare the anguish of his still unconscious wife, Katherine (Julia Stiles), Robert accepts the suggestion of the hospital's Catholic priest, Father Spiletto (Giovanni Lombardo Radice) to secretly adopt another newborn whose mother died in childbirth. Robert and Katherine raise the boy as their own and name him Damien.

Robert's career ascends over the course of the next five years. He is named Deputy Ambassador to the United Kingdom, and after the ambassador dies in a bizarre accident, becomes Ambassador and settles into a large estate just outside London. However, disturbing events start to revolve around Damien (Seamus Davey-Fitzpatrick), the most prominent being his nanny hanging herself at his birthday party, driven to do so by a large black dog.

Soon afterward, Robert is approached by Father Brennan (Pete Postlethwaite), who delivers an eerie warning and seems to know the truth about Damien's birth, at which point Robert has him escorted out. Meanwhile, photographer Keith Jennings (David Thewlis) finds out that several of his photographs contain mysterious signs, later revealed to foreshadow a series of shocking deaths.